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1.
Carbon and nitrogen isotopic analysis of the Middle/Late Bronze individuals from Bezdanjača Cave, Lika region, Croatia
Mario Carić, Emily Zavodny, Mario Novak, Ivor Janković, Siniša Radović, Sarah B. McClure, 2026, original scientific article

Abstract: Almost immediately upon its discovery in 1965, the Middle-Late Bronze Age cave site of Bezdanjača was recognized as one of the earliest, largest, and most important cemeteries in the Lika region of Croatia. Yet, despite its significance, not much is known about the people who were buried there. By combining bioarchaeological and stable carbon and nitrogen isotope analyses of a subset of the burial population, this study aims to (i) systematically reconstruct the diet and health of these individuals, (ii) investigate possible differences in dietary habits within this community based on sex and age, and (iii) contextualize results from Bezdanjača with other Bronze Age assemblages from the surrounding area.
Keywords: Bronze Age, Necropolis, cave, diet, millet, childhood health
Published in RUP: 01.04.2026; Views: 785; Downloads: 30
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Evaluating the effects of managed free-roaming cat populations on prey through stable isotope analysis : a pilot study from British Columbia, Canada
Valentina Martinoia, Renee Ferguson, Peter J. Wolf, Mario Carić, Mario Novak, Shelly Roche, 2025, original scientific article

Abstract: Free-roaming domestic cats (Felis catus) present a major management challenge for animal welfare and biodiversity conservation. Trap-neuter-return (TNR) programs, which include sterilization and return of cats, are increasingly adopted to manage cat populations, often alongside routine food provisioning. However, their effectiveness in reducing cats’ reliance on wild prey remains contested. In this study, we use stable isotope analysis (δ13C, δ15N, δ34S) of cat fur to investigate dietary patterns before and after TNR implementation in the context of concurrent changes in food availability linked to the closure of nearby mink-farming operations. We analyzed samples from 122 cats in a large-scale TNR initiative on a rural property in British Columbia, Canada. These included indoor cats (control), free-roaming cats prior to TNR (Group 1), a subset of Group 1 re-sampled months after food provisioning began (Run 2), and newly sampled cats that had been fed regularly before trapping (Group 2). Local prey and food sources were also analyzed to provide a comparative isotopic baseline. Our results show clear dietary shifts following TNR. Group 1 cats exhibited high isotopic variability and elevated δ15N and δ34S values, consistent with wild prey consumption. In contrast, post-TNR cats showed significantly lower and more homogeneous values, aligning closely with those of indoor, kibble-fed cats. These changes are consistent with a reduced dietary reliance on wildlife and raw mink feed following the combination of TNR with regular provisioning and the cessation of mink operations. These findings demonstrate that regular food provisioning in TNR-managed colonies, particularly when combined with broader environmental changes, can significantly alter cat diets and potentially reduce their dependence on wild prey.
Keywords: trap-neuter-return (TNR) programs, stable isotope analysis, free-roaming cats, conservation
Published in RUP: 05.11.2025; Views: 1170; Downloads: 13
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7.
Multidisciplinary study of human remains from the 3rd century mass grave in the Roman city of Mursa, Croatia
Mario Novak, Orhan Efe Efe Yavuz, Mario Carić, Slavica Filipović, Cosimo Posth, 2025, original scientific article

Abstract: During excavations in 2011, a peculiar archaeological feature representing a mass grave with seven completely preserved skeletons was discovered at the site of the Roman period city of Mursa (modern-day Osijek, Croatia). The archaeological context and direct radiocarbon dating indicate that the bodies were interred during the mid-3rd century CE. Bioarchaeological analysis shows that all seven individuals are adult males exhibiting numerous pathological lesions (e.g., enthesopathies, injuries). Carbon and nitrogen stable isotopes analysis indicates they had a mixed C3/C4-based vegetal diet with limited amounts of terrestrial animal protein and a very limited marine protein consumption. Ancient DNA analysis shows that individuals from the Mursa mass grave had a heterogenous ancestry. None of them show genetic continuity with the preceding local Early Iron Age population. The presented multidisciplinary analyses of the Mursa mass grave strongly suggest that the studied individuals were Roman soldiers, victims of a catastrophic event occurring as the result of the ‘Crisis of the Third Century’, most probably the battle of Mursa from 260 CE.
Keywords: battle, bioarchaeology, ancient DNA, Late Antiquity
Published in RUP: 17.10.2025; Views: 1023; Downloads: 8
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8.
Isotope and archaeobotanical analysis reveal radical changes in mobility, diet and inequalities around 1500 BCE at the core of Europe
Claudio Cavazzuti, Anikó Horváth, Anett Gémes, Kristóf Fülöp, Tamás Szeniczey, János Gábor Tarbay, Ashley McCall, Beatriz Gamarra Rubio, Magdolna Vicze, Annamária Bárány, Mario Novak, 2025, original scientific article

Abstract: The transition from the Middle to the Late Bronze Age (around 1500 BCE) in the Carpathian Basin was parallel by drastic cultural changes in Central-Europe, which strongly influenced the dynamic of prehistoric Europe. The cultural fragmentation of the Middle Bronze Age (2000 − 1500 BCE) Carpathian Basin was followed by a more homogeneous development at the beginning of the Late Bronze Age (1500 − 1300 BCE), with the appearance of the Tumulus culture. In the beginning of this period, the long-used tell-settlements were abandoned, furthermore new pottery styles and metal types appeared. Whether these changes were caused by immigration, or a local adaptation to external influxes, has long been a matter of debate. Our study investigates this transition from the point of view of diet and mobility from several key-sites of Hungary. Our results show (1) low migration rates and a shift of migration trajectories; that (2) the beginning of the systematic consumption of Panicum miliaceum was from 1540 − 1480 BCE; that (3) the decrease of average animal protein intake was parallel by an increase of cereal consumption and a tendency to less unequal diet. Overall, our results shed new light on the dynamics of complex change in Bronze Age Europe.
Keywords: Bronze Age, Hungary, stable isotopes
Published in RUP: 26.08.2025; Views: 952; Downloads: 12
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9.
Analiza skeleta iz jame Oporovina (Lovranska Draga) : magistrsko delo
Zorana Dimković, 2019, master's thesis

Published in RUP: 28.11.2019; Views: 5005; Downloads: 118
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